Who holds it, who is betting against it, and where the big money is moving
Net of large-order buying minus selling, in USD. Over this window: −$5.54M (9 inflow days, 11 outflow days).
Super/Big orders approximate institutional-size trades; Mid/Small approximate retail-size trades. Net buy minus sell value in each bucket, summed over the window.
The five largest holders on file together own 46.5% of shares outstanding.
Reported short interest has fallen from 13.91% to 13.82% of float over the last 6 biweekly reports (30 Jun 2026 to 15 Sep 2026).
| Holder | Type | Period | Share change | New stake |
|---|---|---|---|---|
| Putnam LLC | Mutual Fund | 2026/Q3 | +130,655 | 0.0% |
| Putnam Fiduciary Trust Company | Private Company | 2026/Q3 | +124,492 | 0.0% |
| Nuveen Asset Management LLC | Insurance Company | 2026/Q3 | +347,599 | 0.1% |
| Riverbridge Partners, LLC | Mutual Fund | 2026/Q3 | +152,878 | 0.1% |
| RA Capital Management, L.P. | VC/PE Fund | 2026/Q3 | +18,750,000 | 6.3% |
| Bank of America Corporation | Public Company | 2026/Q3 | +2,794,012 | 1.9% |
| BlackRock | Mutual Fund | 2026/Q3 | +2,434,998 | 5.6% |
| Marshall Wace LLP | Hedge Fund | 2026/Q3 | +2,266,979 | 1.0% |
| The Toronto-Dominion Bank | Public Company | 2026/Q3 | +1,972,381 | 0.7% |
| Jacobs Levy Equity Management Inc | Mutual Fund | 2026/Q3 | +1,868,718 | 0.6% |
| Vanguard Capital Management, LLC | Mutual Fund | 2026/Q3 | +1,114,485 | 3.2% |
| State Street Global Advisors | Mutual Fund | 2026/Q3 | +665,048 | 3.7% |
| Point72 Asset Management, L.P. | Hedge Fund | 2026/Q3 | +595,091 | 0.2% |
| Geode Capital Management, LLC | Mutual Fund | 2026/Q3 | +495,986 | 1.6% |
| Goldman Sachs Asset Management, L.P. | Mutual Fund | 2026/Q3 | +457,097 | 0.5% |
Institutional buy/sell activity by reporting period, largest moves first.
| Date | Insider | Title | Transaction | Shares | Price | Held after |
|---|---|---|---|---|---|---|
| 26 Aug 2026 | Dhavalkumar Patel | Chief Scientific Officer | Derivative Exercise and Sale | -41,158 | $4.11 | 183,352 |
| 25 Aug 2026 | Dhavalkumar Patel | Chief Scientific Officer | Derivative Exercise and Retained | +83,842 | – | 183,352 |
| 5 Jul 2026 | Hans Edgar Bishop | Independent Chairperson | Other Acquisition | +6,297 | – | 5,854,800 |
| 3 Jun 2026 | Robert Taylor Nelsen | Independent Director | Other Derivative Acquisition | +90,000 | $2.96 | 90,000 |
| 3 Jun 2026 | Yuhmin Yang | Independent Director | Other Derivative Acquisition | +90,000 | $2.96 | 90,000 |
| 3 Jun 2026 | Michelle Seitz | Independent Director | Other Derivative Acquisition | +90,000 | $2.96 | 90,000 |
| 3 Jun 2026 | Richard Mulligan | Executive Vice Chairman & Head of Innovation Research | Other Derivative Acquisition | +90,000 | $2.96 | 90,000 |
| 3 Jun 2026 | Hans Edgar Bishop | Independent Chairperson | Other Derivative Acquisition | +90,000 | $2.96 | 90,000 |
| 3 Jun 2026 | Joshua Bilenker | Independent Director | Other Derivative Acquisition | +90,000 | $2.96 | 90,000 |
| 3 Jun 2026 | Alise Reicin | Independent Director | Other Derivative Acquisition | +90,000 | $2.96 | 90,000 |
| 3 Jun 2026 | Robert Rosiello | Director | Other Derivative Acquisition | +90,000 | $2.96 | 90,000 |
| 5 Apr 2026 | Hans Edgar Bishop | Independent Chairperson | Other Acquisition | +7,763 | – | 5,848,503 |
| 23 Mar 2026 | Dhavalkumar Patel | Chief Scientific Officer | Initial Statement of Beneficial | +0 | – | 1,396,250 |
| 8 Mar 2026 | Steven Harr | Chief Executive Officer | Derivative Exercise and Sale | -24,345 | $3.13 | 8,960,751 |
| 8 Mar 2026 | Susan Wyrick | Acting CFO, Treasurer, Principal Accounting Officer and Senior VP of Finance & Accounting | Derivative Exercise and Sale | -4,387 | $3.13 | 196,196 |
| 6 Mar 2026 | Steven Harr | Chief Executive Officer | Derivative Exercise and Retained | +67,321 | – | 8,960,751 |
| 5 Mar 2026 | Susan Wyrick | Acting CFO, Treasurer, Principal Accounting Officer and Senior VP of Finance & Accounting | Derivative Exercise and Retained | +9,279 | – | 196,196 |
| 4 Mar 2026 | Steven Harr | Chief Executive Officer | Other Derivative Acquisition | +200,000 | – | 200,000 |
| 4 Mar 2026 | Steven Harr | Chief Executive Officer | Other Derivative Acquisition | +900,000 | $3.41 | 900,000 |
| 4 Mar 2026 | Susan Wyrick | Acting CFO, Treasurer, Principal Accounting Officer and Senior VP of Finance & Accounting | Other Derivative Acquisition | +45,833 | – | 45,833 |
From Form 4/Form 144 filings. Negative share counts are disposals (sales, gifts, or shares withheld for taxes); positive are acquisitions.
| Report date | Shares short | % of float | Days to cover |
|---|---|---|---|
| 15 Sep 2026 | 41,374,811 | 13.8% | 12.2 |
| 31 Aug 2026 | 41,563,334 | 13.9% | 16.1 |
| 14 Aug 2026 | 42,568,975 | 14.2% | 18.0 |
| 31 Jul 2026 | 42,148,007 | 15.1% | 16.3 |
| 15 Jul 2026 | 39,736,380 | 14.3% | 13.2 |
| 30 Jun 2026 | 38,698,572 | 13.9% | 9.1 |
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